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Humpback whale Temporal range: 20–0 Ma PreꞒ Ꞓ O S D C P T J K Pg N Early Miocene – Recent
Size compared to an average human
Conservation status
Least Concern  (IUCN 3.1)
CITES Appendix I (CITES)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Infraorder:	Cetacea
Family:	Balaenopteridae
Genus:	Megaptera Gray, 1846
Species:	M. novaeangliae
Binomial name
Megaptera novaeangliae Borowski, 1781
Humpback whale range
Synonyms
Balaena gibbosa Erxleben, 1777 B. boops Fabricius, 1780 B. nodosa Bonnaterre, 1789 B. longimana Rudolphi, 1832 Megaptera longimana Gray, 1846 Kyphobalaena longimana Van Beneden, 1861 Megaptera versabilis Cope, 1869

The humpback whale (Megaptera novaeangliae) is a species of baleen whale. It is one of the larger rorqual species, with adults ranging in length from 12–16 m (39–52 ft) and weighing around 25–30 t (28–33 short tons). The humpback has a distinctive body shape, with long pectoral fins and a knobbly head. It is known for breaching and other distinctive surface behaviors, making it popular with whale watchers. Males produce a complex song lasting 10 to 20 minutes, which they repeat for hours at a time. All the males in a group will produce the same song, which is different each season.

Found in oceans and seas around the world, humpback whales typically migrate up to 25,000 km (16,000 mi) each year. They feed in polar waters, and migrate to tropical or subtropical waters to breed and give birth, fasting and living off their fat reserves. Their diet consists mostly of krill and small fish. Humpbacks have a diverse repertoire of feeding methods, including the bubble net technique.

Like other large whales, the humpback was a target for the whaling industry. The species was once hunted to the brink of extinction; its population fell by an estimated 90% before a 1966 moratorium. While numbers have partially recovered to some 80,000 animals worldwide, entanglement in fishing gear, collisions with ships and noise pollution continue to affect the species.

Taxonomy

The humpback was first identified as baleine de la Nouvelle Angleterre by Mathurin Jacques Brisson in his Regnum Animale of 1756. In 1781, Georg Heinrich Borowski described the species, converting Brisson's name to its Latin equivalent, Balaena novaeangliae. In 1804, Lacépède shifted the humpback from the family Balaenidae, renaming it B. jubartes. In 1846, John Edward Gray created the genus Megaptera, classifying the humpback as Megaptera longipinna, but in 1932, Remington Kellogg reverted the species names to use Borowski's novaeangliae. The common name is derived from the curving of their backs when diving. The generic name Megaptera from the Ancient Greek mega- μεγα ("giant") and ptera/ πτερα ("wing"), refers to their large front flippers. The specific name means "New Englander" and was probably given by Brisson due to regular sightings of humpbacks off the coast of New England.

Humpback whales are rorquals, members of the Balaenopteridae family that includes the blue, fin, Bryde's, sei and minke whales. The rorquals are believed to have diverged from the other families of the suborder Mysticeti as long ago as the middle Miocene era. Though clearly related to the giant whales of the genus Balaenoptera, the humpback is the sole member of its genus. Recent DNA sequencing has indicated the humpback is actually more closely related to the fin whale (B. physalus) and possibly the gray (Eschrichtius robustus), than it is to others such as the minke.

Genetic research in mid-2014 by the British Antarctic Survey confirmed that the separate populations in the North Atlantic, North Pacific and Southern Oceans are more distinct than previously thought. Some biologists believe that these should be regarded as separate subspecies and that they are evolving independently.

Description

The humpback whale is perhaps the most recognizable baleen whale. It has a highly narrow rostrum, robust body and proportionally long flippers (around one-third its body length). The dorsal fin is generally small but varies in shape from low and almost non-existent to relatively high and curved. Rounded bumps or tubercles exist on the upper and lower jaw and front edge of the flippers, while the tail fluke has a serrated trailing edge. These are key features distinguishing the species from other large whales. As with other rorquals, the humpback has a series of grooves stretching from the tip of the lower jaw to the navel. They are fewer in number in this species, ranging from 14–35. Humpbacks have 270–400 baleen plates on both sides of the mouth.

The dorsal or upper-side of the animal is generally black while the ventral or underside is black, white and mottled in pigmentation. The varying color patterns and scars on the tail flukes distinguish individual animals. The female has a hemispherical lobe about 15 cm (5.9 in) in diameter in her genital region. This visually distinguishes males and females. The male's penis usually remains hidden in the genital slit.

Size

Fully grown males average 13–14 m (43–46 ft). Females are slightly larger at 15–16 m (49–52 ft); one large recorded specimen was 19 m (62 ft) long and had pectoral fins measuring 6 m (20 ft) each. The largest humpback on record, according to whaling records, was a female killed in the Caribbean; she was 27 m (89 ft) long with a weight of 90 metric tons (99 short tons), although the reliability of these extremely atypical data is impossible to confirm. The largest measured by the scientists of the Discovery Committee were a female and a male at 14.9 and 14.75 m (48.9 and 48.4 ft) respectively, although this was out of a sample size of only 63 whales. Body mass typically is in the range of 25–30 metric tons (28–33 short tons), with large specimens weighing over 40 metric tons (44 short tons).

Behavior and ecology

Humpback whales normally associate in small, unstable groups, though large aggregations form during feeding among among males competing for females. They may interact with other cetacean species, such as right whales, fin whales and bottlenose dolphins. Humpbacks can be highly active at the surface, performing aerial behaviors such as breaching, tail slapping (lobtailing) and flipper slapping. These may serve various functions such as play, communication, parasite removal, and displaying excitement or annoyance. Humpbacks rest at the surface in a horizontal position.

Feeding

Humpback whales are generalist feeders, their main food items being krill and small schooling fish. In the southern hemisphere, the most common krill species eaten is the Antarctic krill, while in other places, the northen krill and various species of Euphausia and Thysanoessa are taken. Fish species consumed include herring, capelin, sand lances and Atlantic mackerel. Feeding takes place from spring to fall.

The humpback has the most diverse hunting repertoire of all baleen whales. Its most inventive technique is known as bubble net feeding; a group of whales dive up to 20 m (66 ft) below the surface and swim in a shrinking circle blowing air from their blowholes creating vertical cylinder-ring of bubbles that captures the prey above them. Humpbacks use two main behaviors to create bubble-netting; upward spirals and double loops. Upward spirals involve the whales blowing continuously as they circle towards the surface, creating a spiral of bubble. Double loops consist of a deep, long loop that corrals the prey followed by lobtailing at the surface and then a smaller loop that serves to make the final capture of the prey. After the nets are created, humpbacks swimming into them with their mouth gaping and ready to swallow. Using network-based diffusion analysis, one study argued that whales learned lobtailing from other whales in the group over a period of 27 years in response to a change in the primary form of prey.

Courtship and reproduction

Mating and breeding takes place during the winter months. Females go through estrus while males reach peak testosterone and sprem levels. Humpback whales are promiscuous, with both sexes having multiple partners. Males may sing to attract females or display dominance to other males. WIth high amount of competitive display behavior among males, the mating system of the humpback has been compared to lek mating. Males will frequently trail both single females and cow-calf pairs. These are known as "escorts" and the male that is closest to the female and doing the most aggressive behavior to the other males is known as the "principal escort" while those fighting for his position are known as "challengers". Other following males that are not directly competing to be next to the female are called "secondary escorts". Aggressive behavior between males include tail slashing, ramming and head-butting.

Gestation in the species is around 11.5 months and females reproduce every two years. Humpback whale births have been rarely observed. One birth witnessed off Madagascar occurred within four minutes. Humpback whales have been known to hybridize with other rorquals; there is a well-documented report of a humpback-blue whale hybrid in the South Pacific. Birthing mostly takes place in mid-winter, usually to a single calf. Calves suckle for up to a year but can feed independantly by six months. Humpback reach sexual maturity at 5–10 years depending on the population.

Song

Both male and female humpback whales vocalize, but only males produce the long, loud, complex "song" for which the species is famous. Each song consists of several sounds in a low register, varying in amplitude and frequency and typically lasting from 10 to 20 minutes. Individuals may sing continuously for more than 24 hours. Cetaceans have no vocal cords, instead, they produce sound via a larynx-like structure found in the throat, the mechanism of which has not been clearly identified. Whales do not have to exhale to produce sound.

Whales within a population sing a single song. All North Atlantic humpbacks sing the same song, while those of the North Pacific sing a different song. Each population's song changes slowly over a period of years without repeating. The songs are culturally transmitted, and have been shown in some cases to spread "horizontally" between neighbouring populations over the course of successive breeding seasons.

Many of the whales observed to approach a singer are other males, often resulting in conflict. Singing may, therefore, be a challenge to other males. Some scientists have hypothesized the song may serve an echolocative function. During the feeding season, humpbacks make unrelated vocalizations for herding fish into their bubble nets. Humpback whales make other sounds to communicate, such as grunts, groans, snorts and barks.

Predation

Visible scars indicate that orcas can prey upon juvenile humpbacks. A 2014 study off Western Australia observed that when available in large numbers, young humpbacks can be attacked and sometimes killed by orcas. Moreover, mothers and (possibly related) adults escort neonates to deter such predation. The suggestion is that when humpbacks suffered near-extinction during the whaling era, orcas turned to other prey, but are now resuming their former practice. There is also evidence that humpback whales will defend against or attack killer whales who are attacking either humpback calves or juveniles as well as members of other species, including other cetaceans and pinnipeds.

The great white shark is another confirmed predator of the humpback whale. In 2020, Marine biologists Dines and Gennari et al., published a documented incident of a group of great white sharks exhibiting pack-like behavior to successfully attack and kill a live adult humpback whale. A second incident regarding great white sharks killing humpback whales was documented off the coast of South Africa. The shark recorded instigating the attack was a female nicknamed "Helen". Working alone, the shark attacked a 33 ft (10 m) emaciated and entangled humpback whale by attacking the whale's tail to cripple and bleed the whale before she managed to drown the whale by biting onto its head and pulling it underwater.

Migration

Migratory patterns and social interactions were explored in the 1960s, and by further studies in 1971. Calambokidis et al. provided the "first quantitative assessment of the migratory structure of humpback whales in the entire North Pacific basin."

Range and habitat

Humpbacks inhabit all major oceans, in a wide band running from the Antarctic ice edge to 81°N latitude. The four global populations are North Pacific, Atlantic, Southern Ocean and Indian Ocean populations. These populations are distinct. Although the species has cosmopolitan distribution and is usually not considered to cross the equator line, seasonal observations at Cape Verde suggest possible interactions among populations from both hemispheres. Aside from the Arabian Sea group, year-round presences have been confirmed among British and Norwegian waters. Parts of wintering grounds around the globe have been poorly studied or being undetected, such as around Pitcairn Islands, Northern Mariana Islands (e.g. Marpi and CK Reefs vicinity to Saipan), Volcano Islands, Pasaleng Bay, Trindade and Martin Vaz, Mauritius, and Aldabra.

Whales were once uncommon in the Mediterranean and the Baltic Seas, but have increased their presence in both waters as global populations have recovered. Recent increases within the Mediterranean basin, including re-sightings, indicate that more whales may migrate into the inland sea in the future, not only for wintering but also for feeding. Humpbacks are also showing signs of re-expanding into former ranges, such as Scotland, Skagerrak and Kattegat, as well as Scandinavian fjords such as Kvænangen, where they had not been observed for decades.

In the North Atlantic, feeding areas range from Scandinavia to New England. Breeding occurs in the Caribbean and Cape Verde. In the South Atlantic and Indian Oceans, whales may breed off Brazil, as well the coasts of central, southern and southeastern Africa (including Madagascar). Whale visits into Gulf of Mexico have been infrequent, but occurred in the gulf historically. In the South Atlantic, about 10% of world population of the species possibly migrate to the Gulf of Guinea. Comparison of songs between those at Cape Lopez and Abrolhos Archipelago indicate that trans-Atlantic mixings between western and southeastern populations occur.

A large population spreads across the Hawaiian Islands every winter, ranging from the island of Hawaii in the south to Kure Atoll in the north. These animals feed in areas ranging from the coast of California to the Bering Sea. Humpbacks were first observed in Hawaiian waters in the mid-19th century and might have gained a dominance over North Pacific right whales, as the right whales were hunted to near-extinction. A smaller portion of the north Pacific group migrates in winter to Bahia Banderas, a large bay on the west coast of Mexico in the Puerto Vallarta resort region.

A 2007 study identified seven individuals then wintering off the Pacific coast of Costa Rica as having traveled from the Antarctic—around 8,300 km (5,200 mi). Identified by their unique tail patterns, these animals made the longest documented mammalian migration. In Australia, two main migratory populations were identified, off the west and east coasts. These two populations are distinct, with only a few females in each generation crossing between the two groups. In Panama and Costa Rica, humpback whales come from both the Southern Hemisphere (July–October with over 2,000 whales) and the Northern Hemisphere (December–March numbering about 300).

Some recolonizing habitats are confirmed, especially in the North and South Atlantic (e.g. English and Irish coasts, English Channel) to coasts in the north such as the North Sea and Wadden Sea, where the first confirmed sighting since 1755 was made in 2003, South Pacific (e.g. New Zealand coasts and Niue), pelagic islands of Chile such as Isla Salas y Gómez and Easter Island, where possibilities of undocumented wintering grounds have been considered, southern fiords of Chile and Peru (e.g. Gulf of Penas, Strait of Magellan, Beagle Channel) and in Asia, such as in the Philippines, Babuyan Islands, Cagayan (the first modern mortality of the species in the nation was in 2007), Calayan and Pasaleng Bay, the Ryukyu Islands, the Volcano Islands in Japan, the Northern Mariana Islands recently again became stable/growing wintering grounds while the Marshall Islands, Vietnamese, Taiwanese and Chinese coasts show slow or no obvious recovery. In 2020, A whale was spotted wandering in the Saint Lawrence River at Montreal and a Migaloo in Australian waters.

In Asia

Whales again migrate off Japanese archipelagos and into the Sea of Japan. Connections between these stocks and whales seen in the Sea of Okhotsk, on Kamchatka coasts and around the Commander Islands have been studied. Historical wintering distributions could have been much wider and more southerly distributed, as whales were seen in areas along the Batanes, Sulu and Celebes Seas including off Palawan, Luzon, Malaysia and Mindanao, with higher densities around today's Cape Eluanbi and Kenting National Park. Unconfirmed sightings have been reported near Borneo in modern times. The first confirmation in modern Taiwan was of a pair off Hualien in 1994, followed by a successful escape from entanglement off Taitung in 1999, and continuous sightings around Orchid Island in 2000. Few/none regularly migrate into Kenting National Park. In addition, despite sightings reported almost annually off Green and Orchid Islands, relatively short stays in these waters indicate recoveries as winter foraging has not occurred. Sightings, including of a cow-calf pair, have occurred along the east coast of Taiwan. Around Hong Kong, two documented sightings were recorded in 2009 and in 2016. One of the first documented sightings within the Yellow Sea was of a group of 3 or 4 individuals, including a cow/calf pair in Changhai County in October, 2015.

Since November 2015, whales gather around Hachijō-jima, far north from the known breeding areas in the Bonin Islands. All breeding activities except for giving births had been confirmed as of January, 2016. That makes Hachijo-jima the northernmost breeding ground in the world, north of breeding grounds such as Amami Ōshima, Midway Island, and Bermuda.

Arabian Sea

A non-migratory population in the Arabian Sea remains there year-round. More typical annual migrations cover up to 25,000 km (16,000 mi), making it one of the most-traveled mammalian species. Genetic studies and visual surveys indicate that the Arabian group is the most isolated of all humpback groups and is the most endangered, numbering possibly fewer than 100 animals. Within the Arabian Sea, Masirah Island and Gulf of Masirah, Halaniyat Islands and Kuria Muria Bay are hot spots for the species.

Whales were historically common in continental and marginal waters such as the Hallaniyat Islands, along Indian coasts, Persian Gulf and Gulf of Aden, and recent migrations into the gulf including by cow-calf pairs. It is unknown whether whales seen in the Red Sea originate in this population, however sightings increased since in 2006 even in the northern part of the sea such as in Gulf of Aqaba. Individuals may reach the Pakistan Maldives, Sri Lanka, or further east. Humpbacks have been considered rather vagrant into Persian Gulf, however new studies indicate more regular presences can be expected.

Origins of whales occurring at Maldives are not clear either from Arabian or south Pacific populations, and overlaps are possible.

Relation to humans

Whaling

Humpback whales were hunted as early as the 18th century. By the 19th century, many nations (particularly the United States) were hunting the animal heavily in the Atlantic Ocean and to a lesser extent in the Indian and Pacific Oceans. The late-19th-century introduction of the explosive harpoon allowed whalers to accelerate their take. This, along with hunting in the Antarctic Ocean beginning in 1904, sharply reduced whale populations. During the 20th century, over 200,000 humpbacks were taken, reducing the global population by over 90%. North Atlantic populations dropped to as low as 700 individuals.

Ban

In 1946, the International Whaling Commission (IWC) was founded to oversee the industry. They imposed hunting regulations and created hunting seasons. To prevent extinction, IWC banned commercial humpback whaling in 1966. By then, the global population had been reduced to around 5,000. Prior to commercial whaling, populations could have reached 125,000. North Pacific kills alone are estimated at 28,000. The Soviet Union deliberately under-recorded its catches; the Soviets reported catching 2,820 between 1947 and 1972, but the true number was over 48,000.

As of 2004, hunting was restricted to a few animals each year off the Caribbean island of Bequia in the nation of St. Vincent and the Grenadines. The take is not believed to threaten the local population. Japan had planned to kill 50 humpbacks in the 2007/08 season under its JARPA II research program. The announcement sparked global protests. After a visit to Tokyo by the IWC chair asking the Japanese for their co-operation in sorting out the differences between pro- and antiwhaling nations on the commission, the Japanese whaling fleet agreed to take no humpback whales during the two years it would take to reach a formal agreement. In 2010, the IWC authorized Greenland's native population to hunt a few humpback whales for the following three years.

In Japan, humpback, minkes, sperm and many other smaller Odontoceti, including critically endangered species such as North Pacific right, western gray and northern fin, have been targets of illegal captures. The hunts use harpoons for dolphin hunts or intentionally drive whales into nets, reporting them as cases of entanglement. Humpback meat can be found in markets. In one case, humpbacks of unknown quantities were illegally hunted in the Exclusive Economic Zones of anti-whaling nations such as off Mexico and South Africa.

Whale-watching

Whale watching is the leisure activity of observing humpbacks in the wild. Participants watch from shore or on touring boats. Humpbacks are generally curious about nearby objects. Some individuals, referred to as "friendlies", approach whale-watching boats closely, often staying under or near the boat for many minutes. Because humpbacks are typically easily approachable, curious, identifiable as individuals and display many behaviors, they have become the mainstay of whale tourism around the world. Hawaii has used the concept of "ecotourism" to benefit from the species without killing them. This business brings in revenue of $20 million per year for the state's economy.

Notable individuals

The Tay whale

In December 1883, a male humpback swam up the Firth of Tay in Scotland, past what was then the whaling port of Dundee. Harpooned during a failed hunt, it was found dead off of Stonehaven a week later. Its carcass was exhibited to the public by a local entrepreneur, John Woods, both locally and then as a touring exhibition that traveled to Edinburgh and London. The whale was dissected by Professor John Struthers, who wrote seven papers on its anatomy and an 1889 monograph on the humpback.

Migaloo

An albino humpback whale that travels up and down the east coast of Australia became famous in local media because of its rare, all-white appearance. Migaloo is the only known Australian all-white specimen and is a true albino. First sighted in 1991, the whale was named for an indigenous Australian word for "white fella". To prevent sightseers approaching dangerously close, the Queensland government decreed a 500-m (1600-ft) exclusion zone around him.

Migaloo was last seen in June 2014 along the coast of Cape Byron in Australia. Migaloo has several physical characteristics that can be identified; his dorsal fin is somewhat hooked, and his tail flukes have a unique shape, with edges that are spiked along the lower trailing side.

Humphrey

In 1985, Humphrey swam into San Francisco Bay and then up the Sacramento River towards Rio Vista. Five years later, Humphrey returned and became stuck on a mudflat in San Francisco Bay immediately north of Sierra Point below the view of onlookers from the upper floors of the Dakin Building. He was twice rescued by the Marine Mammal Center and other concerned groups in California. He was pulled off the mudflat with a large cargo net and the help of the US Coast Guard. Both times, he was successfully guided back to the Pacific Ocean using a "sound net" in which people in a flotilla of boats made unpleasant noises behind the whale by banging on steel pipes, a Japanese fishing technique known as oikami. At the same time, the attractive sounds of humpback whales preparing to feed were broadcast from a boat headed towards the open ocean.

Status

Conservation

The worldwide population is estimated at 84,000 and increasing. Regional estimates are 18,000–20,000 in the North Pacific, 12,000 in the North Atlantic and over 50,000 in the Southern Hemisphere, down from a prewhaling population of 125,000.

The NOAA is committed to protecting and recovering the humpback whale. NOAA enacted vessel speed restrictions which serve to protect the endangered North Atlantic right whale and many other species of whales. They respond to dead, injured, or entangled whales. They also educate whale watchers, tourists, and vessel operators on responsible viewing of the humpback whales. NOAA works to develop methods to reduce vessel strikes and reduce risk of entanglement in fishing gear. The NOAA's work will help reduce the number of humpback whale deaths.

In August 2008, the International Union for Conservation of Nature changed humpbacks' status from vulnerable to least concern, although two subpopulations remain endangered. The United States is considering listing separate humpback populations, so smaller groups, such as North Pacific humpbacks, which are estimated to number 18,000–20,000 animals, might be delisted. This is made difficult by humpbacks’ migrations, which can extend 5,157 miles (8,299 km) from Antarctica to Costa Rica. In Costa Rica, the Ballena Marine National Park is designed for humpback protection. Areas where population data is limited and the species may be at higher risk include the Arabian Sea, the western North Pacific Ocean, the west coast of Africa and parts of Oceania.

The species was listed as vulnerable in 1996 and endangered as recently as 1988. Most monitored stocks have rebounded since the end of commercial whaling. In the North Atlantic stocks are believed to be approaching prehunting levels. However, the species is considered endangered in some countries, including the United States.

United States

A 2008 US Department of Commerce analysis (SPLASH) noted that the many challenges to determining the recovery status included the lack of accurate population estimates, the unexpected complexity of population structures and their migration. The report was based on data collected from 2004 to 2006. At the time, the North Pacific population was some 18,302. The estimate is consistent with a moderate rate of recovery for a depleted population, although it was considered to be a "dramatic increase in abundance" from other post-1960s estimates. By comparison, Calambokidis et al. estimated 9,819, covering 1991–1993. This represents a 4% annual increase in population from 1993 to 2006. The sanctuary provided by US national parks, such as Glacier Bay National Park and Preserve and Cape Hatteras National Seashore, became major factors in population recovery.

Canada

Off the west coast of Canada, the Gwaii Haanas National Marine Conservation Area Reserve covers 3,400 square kilometres (1,300 sq mi). It is "a primary feeding habitat" of the North Pacific population. Their critical habitat overlaps with tanker shipping routes between Canada and its eastern trade partners. In 2005 the North Pacific population was listed as threatened under Canada's Species at Risk Act (SARA). On April 19, 2014 the Department of the Environment recommended an amendment to SARA to downgrade their status off the Pacific coast from "threatened" to "species of special concern". According to Committee on the Status of Endangered Wildlife in Canada (COSEWIC), the North Pacific humpback population increased at about 4% annually from 1992 to 2008. Although socioeconomic costs and benefits were considered in their decision to upgrade their status, according to the University of British Columbia's Marine Mammal Research Consortium's research director, the decision was based on biology, not politics.

Threats

Whaling no longer threatens the species, but individuals are vulnerable to collisions with ships, entanglement in fishing gear and noise pollution. Like other cetaceans, humpbacks can be injured by excessive noise. In the 19th century, two humpback whales were found dead near sites of repeated oceanic sub-bottom blasting, with traumatic injuries and fractures in the ears.

Saxitoxin, a paralytic shellfish poisoning from contaminated mackerel, has been implicated in humpback whale deaths.

Whale researchers along the Atlantic Coast report that there have been more stranded whales with signs of vessel strikes and fishing gear entanglement in recent years than ever before. The NOAA recorded 88 stranded humpback whales between January 2016 and February 2019. This is more than double the number of whales stranded between 2013 and 2016. Because of the increase in stranded whales NOAA declared an unusual mortality event in April 2017. This declaration still stands. Virginia Beach aquarium's stranding response coordinator, Alexander Costidis says the conclusions are that the two causes of these unusual mortality events are vessel interactions and entanglements.

See also

• List of cetaceans
• Wilhelmina Bay
• List of animals with humps

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Sources

Books

• Clapham, Phil (1996). Humpback Whales. Voyageur Press. ISBN 978-0-89658-296-5.
• Clapham, Phil (2002). "Humpback Whale". Encyclopedia of Marine Mammals. San Diego: Academic Press. ISBN 978-0-12-551340-1.
• Dawbin, William H. (1966). The Seasonal Migratory Cycle of Humpback Whales. University of California Press.
• Dawes, John; Campbell, Andrew (2008). Exploring the World of Aquatic Life. Infobase Publishing. ISBN 978-1-60413-255-7.
• Evans, Peter G.H.; Raga, Juan Antonio (6 December 2012). Marine Mammals: Biology and Conservation. Springer Science & Business Media. ISBN 978-1-4615-0529-7.
• Reeves, Randall R.; Stewart, Brent S.; Clapham, Phillip J.; Powell, James A. (2002). National Audubon Society guide to marine mammals of the world. Alfred A. Knopf.
• Struthers, Sir John (1889). Memoir on the Anatomy of the Humpback Whale, Megaptera Longimana. Maclachlan and Stewart.

Journal articles

• Best, P. B. (1993). "Increase rates in severely depleted stocks of baleen whales". ICES Journal of Marine Science. 50 (2): 169–186. doi:10.1006/jmsc.1993.1018.
• Smith, T.D.; Allen, J.; Clapham, P.J.; Hammond, P.S.; Katona, S.; Larsen, F.; Lien, J.; Mattila, D.; Palsboll, P.J.; Sigurjonsson, J.; Stevick, P. T.; Oien, N. (1999). "An ocean-basin-wide mark-recapture study of the North Atlantic humpback whale". Marine Mammal Science. 15: 1–32. doi:10.1111/j.1748-7692.1999.tb00779.x. Archived from the original on 2021-06-23. Retrieved 2019-09-16.
• Franklin, T.; Franklin, W.; Brooks, L.; Harrison, P.; Baverstock, P.; Clapham, P. (2011). "Seasonal changes in pod characteristics of eastern Australian humpback whales (Megaptera novaeangliae), Hervey Bay 1992–2005". Marine Mammal Science. 27 (3): E134 – E152. doi:10.1111/j.1748-7692.2010.00430.x.

External links

General

• US National Marine Fisheries Service Humpback Whale web page
• ARKive – images and movies of the humpback whale (Megaptera novaeangliae).
• Humpbacks of Hervey Bay, Queensland, Australia
• The Dolphin Institute Whale Resource Guide and scientific publications
• Humpback Whale Gallery (Silverbanks)
• (in French) Humpback whale videos
• The Humpback Whales of Hervey Bay
• Epic humpback whale battle filmed

Humpback whale songs

• The Whalesong Project
• Article from PHYSORG.com on the complex syntax of whalesong phrases
• Voices of the Sea – Sounds of the Humpback Whale Archived 2014-08-22 at the Wayback Machine
• Songlines – Songs of the Eastern Australian Humpback whales

Conservation

• The Oceania Project, Humpback Whale Research, Hervey Bay

Videos

• Humpback whales' attempt to stop killer whale attack – Planet Earth Live – BBC One
• Humpback whales defend Gray whale against Killer whales (YouTube)
• Humpbacks Block Orcas’ Feeding Frenzy (LiveScience)
• Humpback whales charge group of transient orcas (Save Our Seas Foundation)
• Humpbacks Chase Killer Whales Right Under Our Boat, 8/24/2014
• Humpback Whale Mother Fights Off Males to Protect Calf | BBC Earth
• Whale Protects Diver From Shark | The Dodo

Other

• Dead calf at the Amazon rainforest

v t e Genera of baleen whales and their extinct allies
Kingdom: Animalia Phylum: Chordata Class: Mammalia Order: Artiodactyla Infraorder: Cetacea
Mysticeti Mysticeti †Borealodon †Coronodon †Imerocetus †Mioceta? †Piscocetus †Siphonocetus? †Tretulias? †Ulias? †Aetiocetidae Aetiocetus Ashorocetus Chonecetus Fucaia Morawanocetus Willungacetus †Llanocetidae Llanocetus Mystacodon †Mammalodontidae Janjucetus Mammalodon Chaeomysticeti Horopeta Sitsqwayk Whakakai †Eomysticetoidea Cetotheriopsis †Eomysticetidae Eomysticetus Micromysticetus Tohoraata Tokarahia Waharoa Yamatocetus Balaenomorpha Balaenidae †Antwerpibalaena †Archaeobalaena Balaena †Balaenella †Balaenula †Balaenotus Eubalaena †Idiocetus †Mesoteras †Morenocetus †Peripolocetus †Protobalaena Thalassotherii †Hibacetus †Isanacetus †Isocetus †Mauicetus †Pinocetus †Taikicetus †Tiphyocetus †Uranocetus †Aglaocetidae Aglaocetus †Diorocetidae Amphicetus Diorocetus Plesiocetopsis Thinocetus †Pelocetidae Cophocetus Halicetus Parietobalaena Pelocetus †Tranatocetidae Mesocetus Mixocetus Tranatocetus Cetotheriidae †Cephalotropis †Journocetus †Otradnocetus †Palaeobalaena? †Titanocetus? †Tiucetus †Cetotheriinae Brandtocetus Cetotherium Ciuciulea Eucetotherium Kurdalagonus Mithridatocetus Vampalus Zygiocetus Herpetocetinae Caperea †Herentalia †Herpetocetus †Metopocetus †Miocaperea †Nannocetus †Piscobalaena Balaenopteroidea †Eobalaenoptera Balaenopteridae †Archaebalaenoptera †Archaeschrichtius Balaenoptera †Burtinopsis? †Cetoteriophanes †Diunatans †Eschrichtioides Eschrichtius †Gricetoides †Incakujira Megaptera †Megapteropsis? †Miobalaenoptera †Nehalaennia †Notiocetus †Parabalaenoptera †Plesiobalaenoptera †Plesiocetus †Praemegaptera †Protororqualus

• IUCN Red List least concern species
• Mammals described in 1781
• Baleen whales
• Cosmopolitan species
• ESA threatened species
• Taxa named by Georg Heinrich Borowski