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{{Short description|Genus of theropod dinosaurs}} {{Short description|Genus of early sauropodomorph dinosaurs}}
{{Speciesbox {{Speciesbox
| fossil_range = ] | fossil_range = ] (]), ~{{fossilrange|230}}
| image = | image = Ahvaytum Skeletal.svg
| image_caption = | image_caption = Skeletal reconstruction
| display_parents = 2 | display_parents = 2
| genus = Ahvaytum | genus = Ahvaytum
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'''''Ahvaytum''''' (meaning "long ago") is an ] genus of probable ] ] ]n dinosaurs from the ] ] of ], United States. The genus contains a ], '''''A. bahndooiveche''''', known from a fragmentary hindlimb bones.<ref name="Ahvaytum">{{Cite journal |last1=Lovelace |first1=David M |last2=Kufner |first2=Aaron M |last3=Fitch |first3=Adam J |last4=Curry Rogers |first4=Kristina |last5=Schmitz |first5=Mark |last6=Schwartz |first6=Darin M |last7=LeClair-Diaz |first7=Amanda |last8=St.Clair |first8=Lynette |last9=Mann |first9=Joshua |last10=Teran |first10=Reba |date=2025-01-01 |title=Rethinking dinosaur origins: oldest known equatorial dinosaur-bearing assemblage (mid-late Carnian Popo Agie FM, Wyoming, USA) |journal=Zoological Journal of the Linnean Society |volume=203 |issue=1 |pages=zlae153 |doi=10.1093/zoolinnean/zlae153 |issn=0024-4082}}</ref> '''''Ahvaytum''''' ({{IPA-all|ɔveɪtəm}} {{respell|ah|VAY|tum}}; {{lit|long ago}}) is an ] genus of probable ] ] ]n dinosaurs from the ] ] of ], United States. The genus contains a ], '''''A. bahndooiveche''''', known from fragmentary hindlimb bones. ''Ahvaytum'' represents the oldest known named dinosaur of the ancient ]n landmass.

== Discovery and naming ==
The ''Ahvaytum'' fossil material was discovered in 2013 in outcrops of the lower ] (Garrett's Surprise locality) in west-central Wyoming, United States.<ref name="Press1">{{Cite web |title=Dinosaurs roamed the northern hemisphere millions of years earlier than previously thought, according to new analysis of the oldest North American fossils |url=https://news.wisc.edu/dinosaurs-roamed-the-northern-hemisphere-millions-of-years-earlier-than-previously-thought-according-to-new-analysis-of-the-oldest-north-american-fossils/ |access-date=2025-01-08 |website=] |language=en-US |last1=Cushman |first1=Will }}</ref><ref name=Lovelace2024>{{cite journal |last1=Lovelace |first1=D.M. |last2=Fitch |first2=A.J. |last3=Schwartz |first3=D. |last4=Schmitz |first4=M. |year=2024 |title=Concurrence of Late Triassic lithostratigraphic, radioisotopic, and biostratigraphic data support a Carnian age for the Popo Agie Formation (Chugwater Group), Wyoming, US |journal=] |volume=136 |issue=5-6 |pages=2305–2324 |doi=10.1130/B36807.1}}</ref> The ] specimen, ] 1975, is an isolated left ]. Specimen UWGM 7549, a partial left ] (] end) was referred to ''Ahvaytum'' since it was found within a {{convert|5|m|ft}} ] of the holotype and its anatomy is consistent with saurischians.<ref name="Ahvaytum">{{Cite journal |last1=Lovelace |first1=David M |last2=Kufner |first2=Aaron M |last3=Fitch |first3=Adam J |last4=Curry Rogers |first4=Kristina |last5=Schmitz |first5=Mark |last6=Schwartz |first6=Darin M |last7=LeClair-Diaz |first7=Amanda |last8=St.Clair |first8=Lynette |last9=Mann |first9=Joshua |last10=Teran |first10=Reba |date=2025-01-01 |title=Rethinking dinosaur origins: oldest known equatorial dinosaur-bearing assemblage (mid-late Carnian Popo Agie FM, Wyoming, USA) |journal=Zoological Journal of the Linnean Society |volume=203 |issue=1 |pages=zlae153 |doi=10.1093/zoolinnean/zlae153 |issn=0024-4082 |url=https://academic.oup.com/zoolinnean/article/203/1/zlae153/7942678}}</ref>

Prior to the formal naming of ''Ahvaytum'', the fossil material was noted in a 2020 ] ], where it was initially interpreted as an early-diverging ] closely related to the ].<ref name="SVP2020">{{Cite conference |last1=Fitch |first1=Adam J. |last2=Lovelace |first2=David M. |last3=Stocker |first3=Michelle R. |date=2020 |title=The oldest dinosaur from the northern hemisphere and the origins of Theropoda |url=https://vertpaleo.org/wp-content/uploads/2021/03/SVP_2020_Program-Abstracts-Volume-FINAL-for-Publishing-1.27.2021.pdf |conference=Society of Vertebrate Paleontology 80th Annual Meeting |pages=140–141}}</ref>

In 2025, Lovelace et al. ] ''Ahvaytum bahndooiveche'' as a new genus and species of early sauropodomorphs based on these fossil remains. The ], ''Ahvaytum'' ({{IPA-all|ɔveɪtəm}}), means "long ago", referring to the specimen's old age. The ], ''bahndooiveche'' ({{IPA-all|bɔnduivitʃi}}), literally translates to "water's young handsome man", and is the term used to refer to both dinosaurs and the colorful native ]s. The full binomial (intended to mean "long ago dinosaur") was created by ] elders and students in their native language, intended to counteract the perceived ] associated with the erection of names derived from European languages.<ref name="Ahvaytum"/>

''Ahvaytum'' notably represents the oldest dinosaur from the ancient landmass of ] (comprising what is now North America, Europe, and Asia excluding the Indian subcontinent and the Arabian peninsula). Prior to its description, it was assumed that sauropodomorph dinosaurs originated in the landmass of ], although ''Ahvaytum'' shows that this clade had already dispersed more widely.<ref name="Ahvaytum"/>

== Description ==
]
''Ahvaytum'' is a small sauropodomorph, estimated to be {{convert|3|ft|m}} long and {{convert|1|ft|m}} tall.<ref name="Press1" /> '']'', a close relative from Argentina, is known from a larger and more complete skeleton with a total body length of around {{convert|1.3|m|ft}}.<ref>{{cite journal | last1 = Sereno | first1 = Paul C. | last2 = Martínez | first2 = Ricardo N. | last3 = Alcober | first3 = Oscar A. | s2cid = 86006363 | year = 2013 | title = Osteology of ''Eoraptor lunensis'' (Dinosauria, Sauropodomorpha). Basal sauropodomorphs and the vertebrate fossil record of the Ischigualasto Formation (Late Triassic: Carnian-Norian) of Argentina | journal = Journal of Vertebrate Paleontology Memoir | volume = 12 | pages = 83–179 | doi = 10.1080/02724634.2013.820113 }}</ref> The fossil material assigned to ''Ahvaytum'' is interpreted as belonging to at least one individual that had grown past the juvenile ontogenetic stage and was still slowly growing.<ref name="Ahvaytum"/>

== Classification ==
]]]
In their ] Lovelace et al. (2025) consistently recovered ''Ahvaytum'' as closely affiliated with the ]n sauropodomorphs '']'' and '']''. Using maximum parsimony, their analysis placed these taxa, along with the African '']'', as members of the family ]. The authors noted that these results, displayed in the ] below, indicate a more inclusive grouping of saturnaliids than traditionally recognized:<ref name="Ahvaytum"/>
{{clade
|{{clade
|1=]
|label2=]
|2={{clade
|1={{clade
|1='']''
|2='']'' }}
|label3=]
|3={{clade
|1={{clade
|1='']''
|label2=]
|2={{clade
|1={{clade
|1='']''
|2='']''
|3='']'' }}
|4={{clade
|1='''''Ahvaytum'''''
|2={{clade
|1='']''
|2={{clade
|1='']''
|2='']'' }} }} }} }} }}
|3={{clade
|1='']''
|2={{clade
|1=]
}} }} }} }} }}|label1=]}}

==Paleoenvironment==
In 2024, Deckman, Lovelace, and Holland suggested that the Popo Agie Formation represents a common river environment, specifically a distributive ] system.<ref>{{cite journal |last1=Deckman |first1=M.E. |last2=Lovelace |first2=D.M. |last3=Holland |first3=S.M. |year=2024 |title=A Reinterpretation of the Jelm and Popo Agie Formations (Triassic, Wyoming) as a Distributive Fluvial System (DFS) and the Role of the Accommodation/Sedimentation Ratio in DFS Deposition |journal=The Mountain Geologist |volume=61 |issue=3 |pages=219–248 |doi=10.31582/rmag.mg.61.3.219}}</ref> The formation's lower unit from which ''Ahvaytum'' is known has also yielded fossils of sulcimentisaurian ']', the large ]s '']'' and '']'' and the ] '']''. The ] '']'', the ]ns '']'' and '']'', and the ] '']'' are known from other layers of this formation.<ref name=Ahvaytum />


== References == == References ==
{{Reflist}} {{Reflist}}


{{Theropoda}} {{Sauropodomorpha}}
{{Taxonbar|from=}} {{Taxonbar|from1=Q131701579|from2=Q131701585}}


] ]
]
]
]
] ]
]

Latest revision as of 23:41, 14 January 2025

Genus of early sauropodomorph dinosaurs

Ahvaytum
Temporal range: Late Triassic (Carnian), ~230 Ma PreꞒ O S D C P T J K Pg N
Skeletal reconstruction
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Clade: Dinosauria
Clade: Saurischia
Clade: Sauropodomorpha (?)
Genus: Ahvaytum
Lovelace et al., 2025
Species: A. bahndooiveche
Binomial name
Ahvaytum bahndooiveche
Lovelace et al., 2025

Ahvaytum (IPA: [ɔveɪtəm] ah-VAY-tum; lit. 'long ago') is an extinct genus of probable basal sauropodomorph saurischian dinosaurs from the Late Triassic Popo Agie Formation of Wyoming, United States. The genus contains a single species, A. bahndooiveche, known from fragmentary hindlimb bones. Ahvaytum represents the oldest known named dinosaur of the ancient Laurasian landmass.

Discovery and naming

The Ahvaytum fossil material was discovered in 2013 in outcrops of the lower Popo Agie Formation (Garrett's Surprise locality) in west-central Wyoming, United States. The holotype specimen, UWGM 1975, is an isolated left astragalus. Specimen UWGM 7549, a partial left femur (proximal end) was referred to Ahvaytum since it was found within a 5 metres (16 ft) radius of the holotype and its anatomy is consistent with saurischians.

Prior to the formal naming of Ahvaytum, the fossil material was noted in a 2020 conference abstracts, where it was initially interpreted as an early-diverging theropod closely related to the Neotheropoda.

In 2025, Lovelace et al. described Ahvaytum bahndooiveche as a new genus and species of early sauropodomorphs based on these fossil remains. The generic name, Ahvaytum (IPA: [ɔveɪtəm]), means "long ago", referring to the specimen's old age. The specific name, bahndooiveche (IPA: [bɔnduivitʃi]), literally translates to "water's young handsome man", and is the term used to refer to both dinosaurs and the colorful native salamanders. The full binomial (intended to mean "long ago dinosaur") was created by Eastern Shoshone elders and students in their native language, intended to counteract the perceived colonialism associated with the erection of names derived from European languages.

Ahvaytum notably represents the oldest dinosaur from the ancient landmass of Laurasia (comprising what is now North America, Europe, and Asia excluding the Indian subcontinent and the Arabian peninsula). Prior to its description, it was assumed that sauropodomorph dinosaurs originated in the landmass of Gondwana, although Ahvaytum shows that this clade had already dispersed more widely.

Description

Size compared to a human

Ahvaytum is a small sauropodomorph, estimated to be 3 feet (0.91 m) long and 1 foot (0.30 m) tall. Eoraptor, a close relative from Argentina, is known from a larger and more complete skeleton with a total body length of around 1.3 metres (4.3 ft). The fossil material assigned to Ahvaytum is interpreted as belonging to at least one individual that had grown past the juvenile ontogenetic stage and was still slowly growing.

Classification

Speculative life restoration

In their phylogenetic analyses Lovelace et al. (2025) consistently recovered Ahvaytum as closely affiliated with the Gondwanan sauropodomorphs Eoraptor and Buriolestes. Using maximum parsimony, their analysis placed these taxa, along with the African Mbiresaurus, as members of the family Saturnaliidae. The authors noted that these results, displayed in the cladogram below, indicate a more inclusive grouping of saturnaliids than traditionally recognized:

Eusaurischia

Theropoda

Sauropodomorpha

Panphagia

Pampadromaeus

Bagualosauria

Bagualosaurus

Saturnaliidae

Saturnalia

Nhandumirim

Chromogisaurus

Ahvaytum

Mbiresaurus

Buriolestes

Eoraptor

Thecodontosaurus

Plateosauria

Paleoenvironment

In 2024, Deckman, Lovelace, and Holland suggested that the Popo Agie Formation represents a common river environment, specifically a distributive fluvial system. The formation's lower unit from which Ahvaytum is known has also yielded fossils of sulcimentisaurian 'silesaurids', the large temnospondyls Anaschisma and Buettnererpeton and the phytosaur Parasuchus. The rhynchosaur Beesiiwo, the pseudosuchians Heptasuchus and Poposaurus, and the dicynodont Eubrachiosaurus are known from other layers of this formation.

References

  1. ^ Cushman, Will. "Dinosaurs roamed the northern hemisphere millions of years earlier than previously thought, according to new analysis of the oldest North American fossils". University of Wisconsin–Madison. Retrieved 2025-01-08.
  2. Lovelace, D.M.; Fitch, A.J.; Schwartz, D.; Schmitz, M. (2024). "Concurrence of Late Triassic lithostratigraphic, radioisotopic, and biostratigraphic data support a Carnian age for the Popo Agie Formation (Chugwater Group), Wyoming, US". GSA Bulletin. 136 (5–6): 2305–2324. doi:10.1130/B36807.1.
  3. ^ Lovelace, David M; Kufner, Aaron M; Fitch, Adam J; Curry Rogers, Kristina; Schmitz, Mark; Schwartz, Darin M; LeClair-Diaz, Amanda; St.Clair, Lynette; Mann, Joshua; Teran, Reba (2025-01-01). "Rethinking dinosaur origins: oldest known equatorial dinosaur-bearing assemblage (mid-late Carnian Popo Agie FM, Wyoming, USA)". Zoological Journal of the Linnean Society. 203 (1): zlae153. doi:10.1093/zoolinnean/zlae153. ISSN 0024-4082.
  4. Fitch, Adam J.; Lovelace, David M.; Stocker, Michelle R. (2020). The oldest dinosaur from the northern hemisphere and the origins of Theropoda (PDF). Society of Vertebrate Paleontology 80th Annual Meeting. pp. 140–141.
  5. Sereno, Paul C.; Martínez, Ricardo N.; Alcober, Oscar A. (2013). "Osteology of Eoraptor lunensis (Dinosauria, Sauropodomorpha). Basal sauropodomorphs and the vertebrate fossil record of the Ischigualasto Formation (Late Triassic: Carnian-Norian) of Argentina". Journal of Vertebrate Paleontology Memoir. 12: 83–179. doi:10.1080/02724634.2013.820113. S2CID 86006363.
  6. Deckman, M.E.; Lovelace, D.M.; Holland, S.M. (2024). "A Reinterpretation of the Jelm and Popo Agie Formations (Triassic, Wyoming) as a Distributive Fluvial System (DFS) and the Role of the Accommodation/Sedimentation Ratio in DFS Deposition". The Mountain Geologist. 61 (3): 219–248. doi:10.31582/rmag.mg.61.3.219.
Sauropodomorpha
Avemetatarsalia
Sauropodomorpha
    • see below↓
Sauropodomorpha
Sauropodomorpha
Saturnaliidae
Bagualosauria
Unaysauridae
Plateosauria
Plateosauridae
Massopoda
Riojasauridae
Massospondylidae
Sauropodiformes
Anchisauria
Sauropoda
    • see below↓
Buriolestes schultzi

Pantydraco caducus Massospondylus carinatus

Jingshanosaurus xinwaensis
Sauropoda
Sauropoda
Lessemsauridae
Gravisauria
Vulcanodontidae
Eusauropoda
Cetiosauridae
Mamenchisauridae
Turiasauria
Neosauropoda
Diplodocoidea
  • (see below ↓ )
Macronaria
  • (see below ↓ )
Dubious sauropods
Vulcanodon karibaensis

Barapasaurus tagorei Patagosaurus fariasi

Turiasaurus riodevnesis
Diplodocoidea
Diplodocoidea
Diplodocimorpha
Rebbachisauridae
Khebbashia
Limaysaurinae
Rebbachisaurinae
Flagellicaudata
Dicraeosauridae
Diplodocidae
Apatosaurinae
Diplodocinae
Dicraeosaurus hansemanni Diplodocus carnegii
Macronaria
Macronaria
Camarasauridae
Titanosauriformes
Brachiosauridae
Somphospondyli
Euhelopodidae
Diamantinasauria
Titanosauria
    • see below↓
Pelorosaurus brevis

Sauroposeidon proteles

Wintonotitan wattsi
Titanosauria
Titanosauria
Lirainosaurinae
Eutitanosauria
Colossosauria
Rinconsauria
Aeolosaurini
Lognkosauria
Saltasauroidea
Nemegtosauridae
Saltasauridae
Opisthocoelicaudiinae
Saltasaurinae
Dubious titanosaurs
Andesaurus delgadoi

Ampelosaurus atacis Futalognkosaurus dukei

Saltasaurus loricatus
Topics in sauropodomorph research
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