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Panthera uncia pyrenaica

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Extinct subspecies of felid

Panthera uncia pyrenaica
Temporal range: 0.57–0.53 Ma PreꞒ O S D C P T J K Pg N
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Suborder: Feliformia
Family: Felidae
Subfamily: Pantherinae
Genus: Panthera
Species: P. uncia
Subspecies: P. u. pyrenaica
Trinomial name
Panthera uncia pyrenaica
Hemmer, 2022
Synonyms

Panthera pardus tautavelensis Testu et al. 2010

Panthera uncia pyrenaica, or Panthera pyrenaica, is an extinct pantherine felid that lived during the Middle Pleistocene in southern France. This taxon was initially suggested to be an European paleosubspecies of the modern snow leopard, but subsequently it has been assigned as a separate species within the genus Panthera.

Taxonomy

The holotype mandible was originally interpreted as a leopard fossil, and given the name Panthera pardus tautavelensis in the Program/Guide book for 16th International Cave Bear and Lion Symposium; however, this name, since it was not formally published in a scientific journal, is considered a nomen nudum. Hemmer who described the specimen in 2022 suggested that it represents a subspecies of the modern snow leopard, so he named the taxon as P. u. pyrenaica and attributed the colloquial names "European snow leopard" and "Arago snow leopard".

However, the 2025 study which described another possible European paleosubspecies (P. u. lusitana) recovered P. u. pyrenaica outside the modern snow leopard as P. pyrenaica due to the lack of similar traits, though it may be a basal related species, and claimed that its primitive traits are likely symplesiomorphic with the modern leopard instead. The same study also assigned an Early Pleistocene specimen from Longdan (Linxia Basin) of China as P. aff. pyrenaica, and the traits observed in both specimens might indicate that this taxon is likely less adapted to the cold environment and hunting caprines on the mountains.

References

  1. ^ Hemmer, H. (2023). "An intriguing find of an early Middle Pleistocene European snow leopard, Panthera uncia pyrenaica ssp. nov. (Mammalia, Carnivora, Felidae), from the Arago cave (Tautavel, Pyrénées-Orientales, France)". Palaeobiodiversity and Palaeoenvironments. 103: 207–220. doi:10.1007/s12549-021-00514-y. S2CID 246433218.
  2. Testu, A.; Moigne, A.-M.; Lumley, H. de (2010). "La panthère Panthera pardus tautavelensis nov. ssp. des niveaux inférieurs de la caune de l'Arago à Tautavel (Pyrenées-Orientales, France) dans le contexte des Felidae (Felinae, Pantherinae) de taille moyenne du Pléistocène européen". 16th International Cave Bear and Lion Symposium, Azé (Saône-et-Loire, France, September 22nd-26th, 2010 (in French). Program/Guide book of excursions: 64.
  3. Jiangzuo, Q.; Madurell-Malapeira, J.; Li, X.; Estraviz-López, D.; Mateus, O.; Testu, A.; Li, S.; Wang, S.; Deng, T. (2025). "Insights on the evolution and adaptation toward high-altitude and cold environments in the snow leopard lineage". Science Advances. 11 (3): eadp5243. doi:10.1126/sciadv.adp5243. PMC 11734717. PMID 39813339.
Extant Carnivora species
Feliformia ("cat-like" carnivorans)
Feliformia
Feloidea
Prionodon (Asiatic linsangs)
Felidae (cats)
Pantherinae
Neofelis
Panthera
Felinae sensu stricto
Bay cat
lineage
Pardofelis
Catopuma
Caracal
lineage
Caracal
Leopardus
Lynx
Puma
lineage
Acinonyx
Puma
Leopard cat
lineage
Prionailurus
Felis
Viverroidea
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Viverroidea
Viverridae
Palm civets
Hemigalinae
Paradoxurinae
Paradoxurus
Viverrinae sensu lato
Viverrinae
sensu stricto
Viverra
Genettinae
Poiana
(African linsangs)
Genetta (genets)
subgenus Genetta
(paraphyletic)
subgenus Eugenetta
(paraphyletic)
subgenus Herpailuropoda
(paraphyletic)
subgenus Pardogale
(paraphyletic)
subgenus Prionailuropoda
subgenus Leptailuropoda
(paraphyletic)
subgenus Osbornictis
Herpestoidea
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Herpestoidea
Hyaenidae
(hyenas)
Proteles
Hyaeninae
(bone-crushing hyenas)
Crocuta
Herpestidae sensu lato
Eupleridae
(Malagasy
carnivorans)
Euplerinae
(Malagasy civets)
Eupleres (falanoucs)
Galidiinae
(vontsira)
Galidictis
Salanoia
Herpestidae sensu stricto (mongooses)
Mungotinae
Suricata
Mungos
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Crossarchus
(kusimanses)
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(Asian mongooses)
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(slender mongooses)
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Canidae (dogs)
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(raccoon dogs)
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(true foxes)
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Cerdocyonina
(zorro)
Speothos
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(South American foxes)
Canina
(wolf-like canids)
Lupulella
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Canis
Arctoidea
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(bears)
Ailuropoda
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Ursinae
Ursus
Mustelida
Pinnipedia (seals)
    • see below↓
Musteloidea
    • see below↓
Pinnipedia (seals)
Odobenidae

Otariidae
(eared seals)
Callorhinus
(northern fur seals)
Otariinae
(sea lions)
Zalophus
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Arctocephalus
(southern fur seals)
Phocidae
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or true seals)
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("northern seals")
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("southern seals")
Monachini
(monk seals)
Neomonachus
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(elephant seals)
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Musteloidea
Ailuridae
Mephitidae
(skunks)
Conepatus
(hog-nosed skunks)
Mephitis
Mydaus
(stink badgers)
Spilogale
(spotted skunks)
Procyonidae
Bassariscus
Procyon
(raccoons)
Bassaricyon
(olingos)
Nasuina
(coatis)
Nasua
Nasuella
(mountain coatis)
Mustelidae
    • see below↓
Mustelidae
Mustelidae
Mellivora
Melinae
Arctonyx
(hog badgers)
Meles
(Eurasian badgers)
Melogale
(ferret-badgers)
Guloninae
Pekania
Gulo
Martes
(martens)
Ictonychinae
Lyncodontini
Galictis
(grisons)
Ictonychini
(African polecats)
Vormela
Ictonyx
Lutrinae (otters)
Lontra
Enhydra
Lutra
Lutrogale
Aonyx
Mustelinae
Neogale
(New World weasels)

Mustela
(weasels)
subgenus Mustela
(paraphyletic)
subgenus Lutreola
(paraphyletic)
subgenus Putorius
Taxon identifiers
Panthera uncia pyrenaica
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